Gas exchange through stomata

Gases enter and leave the leaf through stomata (tiny pores mostly on the lower epidermis). Guard cells control stomatal opening: they open during the day to allow carbon dioxide to enter for photosynthesis, and close at night to reduce water loss by transpiration.

When stomata are open, carbon dioxide diffuses in from the surrounding air because its concentration is higher outside than inside the photosynthesising leaf. Oxygen produced by photosynthesis diffuses out through the same stomata.

Photosynthesis and respiration: the balance

During the day, the rate of photosynthesis exceeds the rate of respiration. The leaf produces more oxygen than it uses and consumes more carbon dioxide than it produces. Net gas exchange: CO₂ in, O₂ out.

At night, only respiration occurs (no light for photosynthesis). The plant uses oxygen and releases carbon dioxide. Net gas exchange: O₂ in, CO₂ out.

At low light levels, there is a compensation point where the rate of photosynthesis exactly equals the rate of respiration. At this point, no net gas exchange with the atmosphere occurs.

Leaf structure for gas exchange

The spongy mesophyll layer has loosely packed cells with large air spaces between them. These air spaces connect to the stomata and provide a large internal surface area for gas exchange. Gases diffuse through the air spaces to reach all mesophyll cells.

The moist cell surfaces of mesophyll cells allow carbon dioxide to dissolve before entering the cells for photosynthesis. The thin cell walls and the short distances between air spaces and cells ensure efficient diffusion throughout the leaf interior.