The two circuits

The pulmonary circuit carries deoxygenated blood from the right ventricle to the lungs via the pulmonary artery. In the lungs, carbon dioxide is released and oxygen is picked up. Oxygenated blood returns to the left atrium via the pulmonary vein.

The systemic circuit carries oxygenated blood from the left ventricle to the body via the aorta. Oxygen and nutrients are delivered to tissues, and carbon dioxide and waste are collected. Deoxygenated blood returns to the right atrium via the vena cava.

Why double circulation is an advantage

In a single circulation (as in fish), blood passes through the heart only once per circuit. After picking up oxygen in the gills, blood flows directly to the body at reduced pressure (because resistance in the gill capillaries slows it). This limits how quickly oxygen can be delivered to active tissues.

In double circulation, oxygenated blood returns to the heart and is pumped a second time to the body. The left ventricle generates high pressure, so blood reaches body tissues quickly and at sufficient pressure to maintain rapid delivery of oxygen and glucose to active cells. This supports the higher metabolic rates of mammals.

Separation of oxygenated and deoxygenated blood

The septum (wall) between the left and right sides of the heart ensures that oxygenated and deoxygenated blood do not mix. The left side handles oxygenated blood; the right side handles deoxygenated blood. This separation maximises the oxygen concentration of blood reaching body tissues.

If the septum has a hole (a septal defect), some mixing occurs, reducing the efficiency of oxygen delivery. This is a congenital heart condition that may require surgical repair.